[1] Univ Nottingham, Plant Sci Div, Loughborough LE12 5RD, Leics, England
[2] Univ Nottingham, Ctr Plant Integrat Biol, Sch Biosci, Loughborough LE12 5RD, Leics, England
[3] Univ Ghent VIB, Dept Plant Syst Biol, B-9052 Ghent, Belgium
[4] Univ Ghent, Dept Plant Biotechnol & Genet, B-9052 Ghent, Belgium
[5] Univ Extremadura, Dept Anat Biol Celular & Zool, E-06071 Badajoz, Spain
[6] Equipe Rhizogenese, UMR DIA PC Agro M INRA IRD UM2, Inst Rech Dev, F-34394 Montpellier 5, France