[1] Univ Dundee, Ninewells Hosp & Med Sch, Dept Med, Ctr Publ Hlth Nutr Res, Dundee DD1 9SY, Scotland
[2] Univ Glasgow, Inst Biomed & Life Sci, Div Biochem & Mol Biol, Glasgow G12 8QQ, Lanark, Scotland
[3] Rowett Res Inst, Greenburn Rd, Bucksburn AB21 9SB, Aberdeen, Scotland