[1] Penn State Univ, Dept Biochem & Mol Biol, HHMI, University Pk, PA 16802 USA
[2] Osaka Univ, Grad Sch Pharmaceut Sci, Lab Environm Biochem, Suita, Osaka 5650871, Japan
[3] SUNY Stony Brook, Dept Biochem & Cell Biol, Stony Brook, NY 11794 USA
[4] Scripps Res Inst, Dept Cell Biol, La Jolla, CA 92037 USA