[1] Univ Queensland, Univ Queensland Ctr Clin Res, Herston, Qld, Australia
[2] Univ Queensland, Queensland Brain Inst, St Lucia, Qld, Australia
[3] Wellcome Trust Sanger Inst, Cambridge, England
[4] Univ Melbourne, Sch Populat Hlth, Ctr Mol Environm Genet & Analyt Epidemiol, Melbourne, Vic 3010, Australia